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The Rise and Progress of Palaeontology

by Thomas Henry Huxley






That application of the sciences of biology and geology; which

is commonly known as palaeontology; took its origin in the mind

of the first person who; finding something like a shell; or a

bone; naturally imbedded in gravel or rock; indulged in

speculations upon the nature of this thing which he had dug out

this 〃fossil〃and upon the causes which had brought it into

such a position。 In this rudimentary form; a high antiquity may

safely be ascribed to palaeontology; inasmuch as we know that;

500 years before the Christian era; the philosophic doctrines of

Xenophanes were influenced by his observations upon the fossil

remains exposed in the quarries of Syracuse。 From this time

forth not only the philosophers; but the poets; the historians;

the geographers of antiquity occasionally refer to fossils;

and; after the revival of learning; lively controversies arose

respecting their real nature。 But hardly more than two centuries

have elapsed since this fundamental problem was first

exhaustively treated; it was only in the last century that the

archaeological value of fossilstheir importance; I mean; as

records of the history of the earthwas fully recognised;

the first adequate investigation of the fossil remains of any

large group of vertebrated animals is to be found in Cuvier's

〃Recherches sur les Ossemens Fossiles;〃 completed in 1822;

and; so modern is stratigraphical palaeontology; that its

founder; William Smith; lived to receive the just recognition of

his services by the award of the first Wollaston Medal in 1831。



But; although palaeontology is a comparatively youthful

scientific speciality; the mass of materials with which it has

to deal is already prodigious。 In the last fifty years the

number of known fossil remains of invertebrated animals has been

trebled or quadrupled。 The work of interpretation of vertebrate

fossils; the foundations of which were so solidly laid by

Cuvier; was carried on; with wonderful vigour and success; by

Agassiz in Switzerland; by Von Meyer in Germany; and last; but

not least; by Owen in this country; while; in later years; a

multitude of workers have laboured in the same field。 In many

groups of the animal kingdom the number of fossil forms already

known is as great as that of the existing species。 In some cases

it is much greater; and there are entire orders of animals of

the existence of which we should know nothing except for the

evidence afforded by fossil remains。 With all this it may be

safely assumed that; at the present moment; we are not

acquainted with a tittle of the fossils which will sooner or

later be discovered。 If we may judge by the profusion yielded

within the last few years by the Tertiary formations of North

America; there seems to be no limit to the multitude of

mammalian remains to be expected from that continent;

and analogy leads us to expect similar riches in Eastern Asia;

whenever the Tertiary formations of that region are as carefully

explored。 Again; we have; as yet; almost everything to learn

respecting the terrestrial population of the Mesozoic epoch;

and it seems as if the Western territories of the United States

were about to prove as instructive in regard to this point as

they have in respect of tertiary life。 My friend Professor Marsh

informs me that; within two years; remains of more than 160

distinct individuals of mammals; belonging to twenty species and

nine genera; have been found in a space not larger than the

floor of a good…sized room; while beds of the same age have

yielded 300 reptiles; varying in size from a length of 60 feet

or 80 feet to the dimensions of a rabbit。



The task which I have set myself to…night is to endeavour to lay

before you; as briefly as possible; a sketch of the successive

steps by which our present knowledge of the facts of

palaeontology and of those conclusions from them which are

indisputable; has been attained; and I beg leave to remind you;

at the outset; that in attempting to sketch the progress of a

branch of knowledge to which innumerable labours have

contributed; my business is rather with generalisations than

with details。 It is my object to mark the epochs of

palaeontology; not to recount all the events of its history。



That which I just now called the fundamental problem of

palaeontology; the question which has to be settled before any

other can be profitably discussed; is this; What is the nature

of fossils? Are they; as the healthy common sense of the ancient

Greeks appears to have led them to assume without hesitation;

the remains of animals and plants? Or are they; as was so

generally maintained in the fifteenth; sixteenth; and

seventeenth centuries; mere figured stones; portions of mineral

matter which have assumed the forms of leaves and shells and

bones; just as those portions of mineral matter which we call

crystals take on the form of regular geometrical solids?

Or; again; are they; as others thought; the products of the

germs of animals and of the seeds of plants which have lost

their way; as it were; in the bowels of the earth; and have

achieved only an imperfect and abortive development? It is easy

to sneer at our ancestors for being disposed to reject the first

in favour of one or other of the last two hypotheses; but it is

much more profitable to try to discover why they; who were

really not one whit less sensible persons than our excellent

selves; should have been led to entertain views which strike us

as absurd; The belief in what is erroneously called spontaneous

generation; that is to say; in the development of living matter

out of mineral matter; apart from the agency of pre…existing

living matter; as an ordinary occurrence at the present day

which is still held by some of us; was universally accepted as

an obvious truth by them。 They could point to the arborescent

forms assumed by hoar…frost and by sundry metallic minerals as

evidence of the existence in nature of a 〃plastic force〃

competent to enable inorganic matter to assume the form of

organised bodies。 Then; as every one who is familiar with

fossils knows; they present innumerable gradations; from shells

and bones which exactly resemble the recent objects; to masses

of mere stone which; however accurately they repeat the outward

form of the organic body; have nothing else in common with it;

and; thence; to mere traces and faint impressions in the

continuous substance of the rock。 What we now know to be the

results of the chemical changes which take place in the course

of fossilisation; by which mineral is substituted for organic

substance; might; in the absence of such knowledge; be fairly

interpreted as the expression of a process of development in the

opposite directionfrom the mineral to the organic。 Moreover;

in an age when it would have seemed the most absurd of paradoxes

to suggest that the general level of the sea is constant; while

that of the solid land fluctuates up and down through thousands

of feet in a secular ground swell; it may well have appeared far

less hazardous to conceive that fossils are sports of nature

than to accept the necessary alternative; that all the inland

regions and highlands; in the rocks of which marine shells had

been found; had once been covered by the ocean。 It is not so

surprising; therefore; as it may at first seem; that although

such men as Leonardo da Vinci and Bernard Palissy took just

views of the nature of fossils; the opinion of the majority of

their contemporaries set strongly the other way; nor even that

error maintained itself long after the scientific grounds of the

true interpretation of fossils had been stated; in a manner that

left nothing to be desired; in the latter half of the

seventeenth century。 The person who rendered this good service

to palaeontology was Nicolas Steno; professor of anatomy in

Florence; though a Dane by birth。 Col

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