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С˵£º the conditions of existence ×ÖÊý£º ÿҳ4000×Ö

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fifty¡¡seeds¡¡shall¡¡grow¡¡up¡¡and¡¡flourish£»¡¡or¡¡whether¡¡it¡¡shall¡¡die¡¡and
perish¡£¡¡¡¡This¡¡is¡¡what¡¡Mr¡£¡¡Darwin¡¡has¡¡drawn¡¡attention¡¡to£»¡¡and¡¡called¡¡the
¡¨STRUGGLE¡¡FOR¡¡EXISTENCE¡¨£»¡¡and¡¡I¡¡have¡¡taken¡¡this¡¡simple¡¡case¡¡of¡¡a¡¡plant
because¡¡some¡¡people¡¡imagine¡¡that¡¡the¡¡phrase¡¡seems¡¡to¡¡imply¡¡a¡¡sort¡¡of
fight¡£

I¡¡have¡¡taken¡¡this¡¡plant¡¡and¡¡shown¡¡you¡¡that¡¡this¡¡is¡¡the¡¡result¡¡of¡¡the
ratio¡¡of¡¡the¡¡increase£»¡¡the¡¡necessary¡¡result¡¡of¡¡the¡¡arrival¡¡of¡¡a¡¡time
coming¡¡for¡¡every¡¡species¡¡when¡¡exactly¡¡as¡¡many¡¡members¡¡must¡¡be¡¡destroyed
as¡¡are¡¡born£»¡¡that¡¡is¡¡the¡¡inevitable¡¡ultimate¡¡result¡¡of¡¡the¡¡rate¡¡of
production¡£¡¡¡¡Now£»¡¡what¡¡is¡¡the¡¡result¡¡of¡¡all¡¡this£¿¡¡¡¡I¡¡have¡¡said¡¡that
there¡¡are¡¡forty¡­nine¡¡struggling¡¡against¡¡every¡¡one£»¡¡and¡¡it¡¡amounts¡¡to
this£»¡¡that¡¡the¡¡smallest¡¡possible¡¡start¡¡given¡¡to¡¡any¡¡one¡¡seed¡¡may¡¡give
it¡¡an¡¡advantage¡¡which¡¡will¡¡enable¡¡it¡¡to¡¡get¡¡ahead¡¡of¡¡all¡¡the¡¡others£»
anything¡¡that¡¡will¡¡enable¡¡any¡¡one¡¡of¡¡these¡¡seeds¡¡to¡¡germinate¡¡six¡¡hours
before¡¡any¡¡of¡¡the¡¡others¡¡will£»¡¡other¡¡things¡¡being¡¡alike£»¡¡enable¡¡it¡¡to
choke¡¡them¡¡out¡¡altogether¡£¡¡¡¡I¡¡have¡¡shown¡¡you¡¡that¡¡there¡¡is¡¡no
particular¡¡in¡¡which¡¡plants¡¡will¡¡not¡¡vary¡¡from¡¡each¡¡other£»¡¡it¡¡is¡¡quite
possible¡¡that¡¡one¡¡of¡¡our¡¡imaginary¡¡plants¡¡may¡¡vary¡¡in¡¡such¡¡a¡¡character
as¡¡the¡¡thickness¡¡of¡¡the¡¡integument¡¡of¡¡its¡¡seeds£»¡¡it¡¡might¡¡happen¡¡that
one¡¡of¡¡the¡¡plants¡¡might¡¡produce¡¡seeds¡¡having¡¡a¡¡thinner¡¡integument£»¡¡and
that¡¡would¡¡enable¡¡the¡¡seeds¡¡of¡¡that¡¡plant¡¡to¡¡germinate¡¡a¡¡little¡¡quicker
than¡¡those¡¡of¡¡any¡¡of¡¡the¡¡others£»¡¡and¡¡those¡¡seeds¡¡would¡¡most¡¡inevitably
extinguish¡¡the¡¡forty¡­nine¡¡times¡¡as¡¡many¡¡that¡¡were¡¡struggling¡¡with¡¡them¡£

I¡¡have¡¡put¡¡it¡¡in¡¡this¡¡way£»¡¡but¡¡you¡¡see¡¡the¡¡practical¡¡result¡¡of¡¡the
process¡¡is¡¡the¡¡same¡¡as¡¡if¡¡some¡¡person¡¡had¡¡nurtured¡¡the¡¡one¡¡and
destroyed¡¡the¡¡other¡¡seeds¡£¡¡¡¡It¡¡does¡¡not¡¡matter¡¡how¡¡the¡¡variation¡¡is
produced£»¡¡so¡¡long¡¡as¡¡it¡¡is¡¡once¡¡allowed¡¡to¡¡occur¡£¡¡¡¡The¡¡variation¡¡in¡¡the
plant¡¡once¡¡fairly¡¡started¡¡tends¡¡to¡¡become¡¡hereditary¡¡and¡¡reproduce
itself£»¡¡the¡¡seeds¡¡would¡¡spread¡¡themselves¡¡in¡¡the¡¡same¡¡way¡¡and¡¡take¡¡part
in¡¡the¡¡struggle¡¡with¡¡the¡¡forty¡­nine¡¡hundred£»¡¡or¡¡forty¡­nine¡¡thousand£»
with¡¡which¡¡they¡¡might¡¡be¡¡exposed¡£¡¡¡¡Thus£»¡¡by¡¡degrees£»¡¡this¡¡variety£»¡¡with
some¡¡slight¡¡organic¡¡change¡¡or¡¡modification£»¡¡must¡¡spread¡¡itself¡¡over¡¡the
whole¡¡surface¡¡of¡¡the¡¡habitable¡¡globe£»¡¡and¡¡extirpate¡¡or¡¡replace¡¡the
other¡¡kinds¡£¡¡¡¡That¡¡is¡¡what¡¡is¡¡meant¡¡by¡¡NATURAL¡¡SELECTION£»¡¡that¡¡is¡¡the
kind¡¡of¡¡argument¡¡by¡¡which¡¡it¡¡is¡¡perfectly¡¡demonstrable¡¡that¡¡the
conditions¡¡of¡¡existence¡¡may¡¡play¡¡exactly¡¡the¡¡same¡¡part¡¡for¡¡natural
varieties¡¡as¡¡man¡¡does¡¡for¡¡domesticated¡¡varieties¡£¡¡¡¡No¡¡one¡¡doubts¡¡at¡¡all
that¡¡particular¡¡circumstances¡¡may¡¡be¡¡more¡¡favourable¡¡for¡¡one¡¡plant¡¡and
less¡¡so¡¡for¡¡another£»¡¡and¡¡the¡¡moment¡¡you¡¡admit¡¡that£»¡¡you¡¡admit¡¡the
selective¡¡power¡¡of¡¡nature¡£¡¡¡¡Now£»¡¡although¡¡I¡¡have¡¡been¡¡putting¡¡a
hypothetical¡¡case£»¡¡you¡¡must¡¡not¡¡suppose¡¡that¡¡I¡¡have¡¡been¡¡reasoning
hypothetically¡£¡¡¡¡There¡¡are¡¡plenty¡¡of¡¡direct¡¡experiments¡¡which¡¡bear¡¡out
what¡¡we¡¡may¡¡call¡¡the¡¡theory¡¡of¡¡natural¡¡selection£»¡¡there¡¡is¡¡extremely
good¡¡authority¡¡for¡¡the¡¡statement¡¡that¡¡if¡¡you¡¡take¡¡the¡¡seed¡¡of¡¡mixed
varieties¡¡of¡¡wheat¡¡and¡¡sow¡¡it£»¡¡collecting¡¡the¡¡seed¡¡next¡¡year¡¡and¡¡sowing
it¡¡again£»¡¡at¡¡length¡¡you¡¡will¡¡find¡¡that¡¡out¡¡of¡¡all¡¡your¡¡varieties¡¡only
two¡¡or¡¡three¡¡have¡¡lived£»¡¡or¡¡perhaps¡¡even¡¡only¡¡one¡£¡¡¡¡There¡¡were¡¡one¡¡or
two¡¡varieties¡¡which¡¡were¡¡best¡¡fitted¡¡to¡¡get¡¡on£»¡¡and¡¡they¡¡have¡¡killed
out¡¡the¡¡other¡¡kinds¡¡in¡¡just¡¡the¡¡same¡¡way¡¡and¡¡with¡¡just¡¡the¡¡same
certainty¡¡as¡¡if¡¡you¡¡had¡¡taken¡¡the¡¡trouble¡¡to¡¡remove¡¡them¡£¡¡¡¡As¡¡I¡¡have
already¡¡said£»¡¡the¡¡operation¡¡of¡¡nature¡¡is¡¡exactly¡¡the¡¡same¡¡as¡¡the
artificial¡¡operation¡¡of¡¡man¡£

But¡¡if¡¡this¡¡be¡¡true¡¡of¡¡that¡¡simple¡¡case£»¡¡which¡¡I¡¡put¡¡before¡¡you£»¡¡where
there¡¡is¡¡nothing¡¡but¡¡the¡¡rivalry¡¡of¡¡one¡¡member¡¡of¡¡a¡¡species¡¡with
others£»¡¡what¡¡must¡¡be¡¡the¡¡operation¡¡of¡¡selective¡¡conditions£»¡¡when¡¡you
recollect¡¡as¡¡a¡¡matter¡¡of¡¡fact£»¡¡that¡¡for¡¡every¡¡species¡¡of¡¡animal¡¡or
plant¡¡there¡¡are¡¡fifty¡¡or¡¡a¡¡hundred¡¡species¡¡which¡¡might¡¡all£»¡¡more¡¡or
less£»¡¡be¡¡comprehended¡¡in¡¡the¡¡same¡¡climate£»¡¡food£»¡¡and¡¡station£»that
every¡¡plant¡¡has¡¡multitudinous¡¡animals¡¡which¡¡prey¡¡upon¡¡it£»¡¡and¡¡which¡¡are
its¡¡direct¡¡opponents£»¡¡and¡¡that¡¡these¡¡have¡¡other¡¡animals¡¡preying¡¡upon
them£»that¡¡every¡¡plant¡¡has¡¡its¡¡indirect¡¡helpers¡¡in¡¡the¡¡birds¡¡that
scatter¡¡abroad¡¡its¡¡seed£»¡¡and¡¡the¡¡animals¡¡that¡¡manure¡¡it¡¡with¡¡their
dung£»I¡¡say£»¡¡when¡¡these¡¡things¡¡are¡¡considered£»¡¡it¡¡seems¡¡impossible
that¡¡any¡¡variation¡¡which¡¡may¡¡arise¡¡in¡¡a¡¡species¡¡in¡¡nature¡¡should¡¡not
tend¡¡in¡¡some¡¡way¡¡or¡¡other¡¡either¡¡to¡¡be¡¡a¡¡little¡¡better¡¡or¡¡worse¡¡than
the¡¡previous¡¡stock£»¡¡if¡¡it¡¡is¡¡a¡¡little¡¡better¡¡it¡¡will¡¡have¡¡an¡¡advantage
over¡¡and¡¡tend¡¡to¡¡extirpate¡¡the¡¡latter¡¡in¡¡this¡¡crush¡¡and¡¡struggle£»¡¡and¡¡if
it¡¡is¡¡a¡¡little¡¡worse¡¡it¡¡will¡¡itself¡¡be¡¡extirpated¡£

I¡¡know¡¡nothing¡¡that¡¡more¡¡appropriately¡¡expresses¡¡this£»¡¡than¡¡the¡¡phrase£»
¡¨the¡¡struggle¡¡for¡¡existence¡¨£»¡¡because¡¡it¡¡brings¡¡before¡¡your¡¡minds£»¡¡in¡¡a
vivid¡¡sort¡¡of¡¡way£»¡¡some¡¡of¡¡the¡¡simplest¡¡possible¡¡circumstances
connected¡¡with¡¡it¡£¡¡¡¡When¡¡a¡¡struggle¡¡is¡¡intense¡¡there¡¡must¡¡be¡¡some¡¡who
are¡¡sure¡¡to¡¡be¡¡trodden¡¡down£»¡¡crushed£»¡¡and¡¡overpowered¡¡by¡¡others£»¡¡and
there¡¡will¡¡be¡¡some¡¡who¡¡just¡¡manage¡¡to¡¡get¡¡through¡¡only¡¡by¡¡the¡¡help¡¡of
the¡¡slightest¡¡accident¡£¡¡¡¡I¡¡recollect¡¡reading¡¡an¡¡account¡¡of¡¡the¡¡famous
retreat¡¡of¡¡the¡¡French¡¡troops£»¡¡under¡¡Napoleon£»¡¡from¡¡Moscow¡£¡¡¡¡Worn¡¡out£»
tired£»¡¡and¡¡dejected£»¡¡they¡¡at¡¡length¡¡came¡¡to¡¡a¡¡great¡¡river¡¡over¡¡which
there¡¡was¡¡but¡¡one¡¡bridge¡¡for¡¡the¡¡passage¡¡of¡¡the¡¡vast¡¡army¡£¡¡Disorganised
and¡¡demoralised¡¡as¡¡that¡¡army¡¡was£»¡¡the¡¡struggle¡¡must¡¡certainly¡¡have¡¡been
a¡¡terrible¡¡oneevery¡¡one¡¡heeding¡¡only¡¡himself£»¡¡and¡¡crushing¡¡through
the¡¡ranks¡¡and¡¡treading¡¡down¡¡his¡¡fellows¡£¡¡¡¡The¡¡writer¡¡of¡¡the¡¡narrative£»
who¡¡was¡¡himself¡¡one¡¡of¡¡those¡¡who¡¡were¡¡fortunate¡¡enough¡¡to¡¡succeed¡¡in
getting¡¡over£»¡¡and¡¡not¡¡among¡¡the¡¡thousands¡¡who¡¡were¡¡left¡¡behind¡¡or
forced¡¡into¡¡the¡¡river£»¡¡ascribed¡¡his¡¡escape¡¡to¡¡the¡¡fact¡¡that¡¡he¡¡saw
striding¡¡onward¡¡through¡¡the¡¡mass¡¡a¡¡great¡¡strong¡¡fellow£»one¡¡of¡¡the
French¡¡Cuirassiers£»¡¡who¡¡had¡¡on¡¡a¡¡large¡¡blue¡¡cloakand¡¡he¡¡had¡¡enough
presence¡¡of¡¡mind¡¡to¡¡catch¡¡and¡¡retain¡¡a¡¡hold¡¡of¡¡this¡¡strong¡¡man's
cloak¡£¡¡¡¡He¡¡says£»¡¡¡¨I¡¡caught¡¡hold¡¡of¡¡his¡¡cloak£»¡¡and¡¡although¡¡he¡¡swore¡¡at
me¡¡and¡¡cut¡¡at¡¡and¡¡struck¡¡me¡¡by¡¡turns£»¡¡and¡¡at¡¡last£»¡¡when¡¡he¡¡found¡¡he
could¡¡not¡¡shake¡¡me¡¡off£»¡¡fell¡¡to¡¡entreating¡¡me¡¡to¡¡leave¡¡go¡¡or¡¡I¡¡should
prevent¡¡him¡¡from¡¡escaping£»¡¡besides¡¡not¡¡assisting¡¡myself£»¡¡I¡¡still¡¡kept
tight¡¡hold¡¡of¡¡him£»¡¡and¡¡would¡¡not¡¡quit¡¡my¡¡grasp¡¡until¡¡he¡¡had¡¡at¡¡last
dragged¡¡me¡¡through¡£¡¨¡¡¡¡Here¡¡you¡¡see¡¡was¡¡a¡¡case¡¡of¡¡selective¡¡savingif
we¡¡may¡¡so¡¡term¡¡itdepending¡¡for¡¡its¡¡success¡¡on¡¡the¡¡strength¡¡of¡¡the
cloth¡¡of¡¡the¡¡Cuirassier's¡¡cloak¡£¡¡¡¡It¡¡is¡¡the¡¡same¡¡in¡¡nature£»¡¡every
species¡¡has¡¡its¡¡bridge¡¡of¡¡Beresina£»¡¡it¡¡has¡¡to¡¡fight¡¡its¡¡way¡¡through¡¡and
struggle¡¡with¡¡other¡¡species£»¡¡and¡¡when¡¡well¡¡nigh¡¡overpowered£»¡¡it¡¡may¡¡be
that¡¡the¡¡smallest¡¡chance£»¡¡something¡¡in¡¡its¡¡colour£»¡¡perhapsthe
minutest¡¡circumstancewill¡¡turn¡¡the¡¡scale¡¡one¡¡way¡¡or¡¡the¡¡other¡£

Suppose¡¡that¡¡by¡¡a¡¡variation¡¡of¡¡the¡¡black¡¡race¡¡it¡¡had¡¡produced¡¡the¡¡white
man¡¡at¡¡any¡¡timeyou¡¡know¡¡that¡¡the¡¡Negroes¡¡are¡¡said¡¡to¡¡believe¡¡this¡¡to
have¡¡been¡¡the¡¡case£»¡¡and¡¡to¡¡imagine¡¡that¡¡Cain¡¡was¡¡the¡¡first¡¡white¡¡man£»
and¡¡that¡¡we¡¡are¡¡his¡¡descendantssuppose¡¡that¡¡this¡¡had¡¡ever¡¡happened£»
and¡¡that¡¡the¡¡first¡¡residence¡¡of¡¡this¡¡human¡¡being¡¡was¡¡on¡¡the¡¡West¡¡Coast
of¡¡Africa¡£¡¡¡¡There¡¡is¡¡no¡¡great¡¡structural¡¡difference¡¡between¡¡the¡¡white
man¡¡and¡¡the¡¡Negro£»¡¡and¡¡yet¡¡there¡¡is¡¡something¡¡so¡¡singularly¡¡different
in¡¡the¡¡constitution¡¡of¡¡the¡¡two£»¡¡that¡¡the¡¡malarias¡¡of¡¡that¡¡country£»¡¡which
do¡¡not¡¡hurt¡¡the¡¡black¡¡at¡¡all£»¡¡cut¡¡off¡¡and¡¡destroy¡¡the¡¡white¡£¡¡¡¡Then¡¡you
see¡¡there¡¡would¡¡have¡¡been¡¡a¡¡selective¡¡operation¡¡performed£»¡¡if¡¡the¡¡white
man¡¡had¡¡risen¡¡in¡¡that¡¡way£»¡¡he¡¡would¡¡have¡¡been¡¡selected¡¡out¡¡and¡¡removed
by¡¡means¡¡of¡¡the¡¡malaria¡£¡¡¡¡Now¡¡there¡¡really¡¡is¡¡a¡¡very¡¡curious¡¡case¡¡of
selection¡¡of¡¡this¡¡sort¡¡among¡¡pigs£»¡¡and¡¡it¡¡is¡¡a¡¡case¡¡of¡¡selection¡¡of
colour¡¡too¡£¡¡¡¡In¡¡the¡¡woods¡¡of¡¡Florida¡¡there¡¡are¡¡a¡¡great¡¡many¡¡pigs£»¡¡and
it¡¡is¡¡a¡¡very¡¡curious¡¡thing¡¡that¡¡they¡¡are¡¡all¡¡black£»¡¡every¡¡one¡¡of¡¡them¡£
Professor¡¡Wyman¡¡was¡¡there¡¡some¡¡years¡¡ago£»¡¡and¡¡on¡¡noticing¡¡no¡¡pigs¡¡but
these¡¡black¡¡ones£»¡¡he¡¡asked¡¡some¡¡of¡¡the¡¡people¡¡how¡¡it¡¡was¡¡that¡¡they¡¡had
no¡¡white¡¡pigs£»¡¡and¡¡the¡¡reply¡¡was¡¡that¡¡in¡¡the¡¡woods¡¡of¡¡Florida¡¡there¡¡was
a¡¡root¡¡which¡¡they¡¡called¡¡the¡¡Paint¡¡Root£»¡¡and¡¡that¡¡if¡¡the¡¡white¡¡pigs
were¡¡to¡¡eat¡¡any¡¡of¡¡it£»¡¡it¡¡had¡¡the¡¡effect¡¡of¡¡making¡¡their¡¡hoofs¡¡crack£»
and¡¡they¡¡died£»¡¡but¡¡if¡¡the¡¡black¡¡pigs¡¡eat¡¡any¡¡of¡¡it£»¡¡it¡¡did¡¡not¡¡hurt
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