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第7章

lectures on evolution-第7章

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were a recent fish。 But the genus Beryx is represented;

at the present day; by very closely allied species which are

living in the Pacific and Atlantic Oceans。 We may go still

farther back。 I have already referred to the fact that the

Carboniferous formations; in Europe and in America; contain the

remains of scorpions in an admirable state of preservation; and

that those scorpions are hardly distinguishable from such as now

live。 I do not mean to say that they are not different; but

close scrutiny is needed in order to distinguish them from

modern scorpions。



More than this。 At the very bottom of the Silurian series; in

beds which are by some authorities referred to the Cambrian

formation; where the signs of life begin to fail useven there;

among the few and scanty animal remains which are discoverable;

we find species of molluscous animals which are so closely

allied to existing forms that; at one time; they were grouped

under the same generic name。 I refer to the well…known 

Lingula of the Lingula flags; lately; in

consequence of some slight differences; placed in the new genus

Lingulella。 Practically; it belongs to the same great

generic group as the Lingula; which is to be found at the

present day upon your own shores and those of many other parts

of the world。



The same truth is exemplified if we turn to certain great

periods of the earth's historyas; for example; the Mesozoic

epoch。 There are groups of reptiles; such as the

Ichthyosauria and the Plesiosauria; which appear

shortly after the commencement of this epoch; and they occur in

vast numbers。 They disappear with the chalk and; throughout the

whole of the great series of Mesozoic rocks; they present no

such modifications as can safely be considered evidence of

progressive modification。



Facts of this kind are undoubtedly fatal to any form of the

doctrine of evolution which postulates the supposition that

there is an intrinsic necessity; on the part of animal forms

which have once come into existence; to undergo continual

modification; and they are as distinctly opposed to any view

which involves the belief; that such modification may occur;

must take place; at the same rate; in all the different types of

animal or vegetable life。 The facts; as I have placed them

before you; obviously directly contradict any form of the

hypothesis of evolution which stands in need of these

two postulates。



But; one great service that has been rendered by Mr。 Darwin to

the doctrine of evolution in general is this: he has shown that

there are two chief factors in the process of evolution: one of

them is the tendency to vary; the existence of which in all

living forms may be proved by observation; the other is the

influence of surrounding conditions upon what I may call the

parent form and the variations which are thus evolved from it。

The cause of the production of variations is a matter not at all

properly understood at present。 Whether variation depends upon

some intricate machineryif I may use the phraseof the living

organism itself; or whether it arises through the influence of

conditions upon that form; is not certain; and the question may;

for the present; be left open。 But the important point is that;

granting the existence of the tendency to the production of

variations; then; whether the variations which are produced

shall survive and supplant the parent; or whether the parent

form shall survive and supplant the variations; is a matter

which depends entirely on those conditions which give rise to

the struggle for existence。 If the surrounding conditions are

such that the parent form is more competent to deal with them;

and flourish in them than the derived forms; then; in the

struggle for existence; the parent form will maintain itself and

the derived forms will be exterminated。 But if; on the contrary;

the conditions are such as to be more favourable to a derived

than to the parent form; the parent form will be extirpated and

the derived form will take its place。 In the first case; there

will be no progression; no change of structure; through any

imaginable series of ages; in the second place there will be

modification of change and form。



Thus the existence of these persistent types; as I have termed

them; is no real obstacle in the way of the theory of evolution。

Take the case of the scorpions to which I have just referred。

No doubt; since the Carboniferous epoch; conditions have always

obtained; such as existed when the scorpions of that epoch

flourished; conditions in which scorpions find themselves better

off; more competent to deal with the difficulties in their way;

than any variation from the scorpion type which they may have

produced; and; for that reason; the scorpion type has persisted;

and has not been supplanted by any other form。 And there is no

reason; in the nature of things; why; as long as this world

exists; if there be conditions more favourable to scorpions than

to any variation which may arise from them; these forms of life

should not persist。



Therefore; the stock objection to the hypothesis of evolution;

based on the long duration of certain animal and vegetable

types; is no objection at all。 The facts of this characterand

they are numerousbelong to that class of evidence which I have

called indifferent。 That is to say; they may afford no direct

support to the doctrine of evolution; but they are capable of

being interpreted in perfect consistency with it。



There is another order of facts belonging to the class of

negative or indifferent evidence。 The great group of Lizards;

which abound in the present world; extends through the whole

series of formations as far back as the Permian; or latest

Palaeozoic; epoch。 These Permian lizards differ astonishingly

little from the lizards which exist at the present day。

Comparing the amount of the differences between them and modern

lizards; with the prodigious lapse of time between the Permian

epoch and the present day; it may be said that the amount of

change is insignificant。 But; when we carry our researches

farther back in time; we find no trace of lizards; nor of any

true reptile whatever; in the whole mass of formations beneath

the Permian。



Now; it is perfectly clear that if our palaeontological

collections are to be taken; even approximately; as an adequate

representation of all the forms of animals and plants that have

ever lived; and if the record furnished by the known series of

beds of stratified rock covers the whole series of events which

constitute the history of life on the globe; such a fact as this

directly contravenes the hypothesis of evolution; because this

hypothesis postulates that the existence of every form must have

been preceded by that of some form little different from it。

Here; however; we have to take into consideration that important

truth so well insisted upon by Lyell and by Darwinthe

imperfection of the geological record。 It can be demonstrated

that the geological record must be incomplete; that it can only

preserve remains found in certain favourable localities and

under particular conditions; that it must be destroyed by

processes of denudation; and obliterated by processes of

metamorphosis。 Beds of rock of any thickness crammed full of

organic remains; may yet; either by the percolation of water

through them; or by the influence of subterranean heat; lose all

trace of these remains; and present the appearance of beds of

rock formed under conditions in which living forms were absent。

Such metamorphic rocks occur in formations of all ages; and; in

various cases; there are very good grounds for the belief that

they have contained organic remains; and that those remains have

been absolutely obliterated。



I insist upon the defects of the geological record the more

because those who have not attended to these matters are apt t

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