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第15章

lectures on evolution-第15章

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peculiarities of that formpeculiarities which tend to show

that the European Hipparion is rather a member of a

collateral branch; than a form in the direct line of succession。

Next; in the backward order in time; is the Miohippus;

which corresponds pretty nearly with the Anchitherium of

Europe。 It presents three complete toesone large median and

two smaller lateral ones; and there is a rudiment of that digit;

which answers to the little finger of the human hand。



The European record of the pedigree of the horse stops here;

in the American Tertiaries; on the contrary; the series of

ancestral equine forms is continued into the Eocene formations。

An older Miocene form; termed Mesohippus; has three toes

in front; with a large splint…like rudiment representing the

little finger; and three toes behind。 The radius and ulna; the

tibia and the fibula; are distinct; and the short crowned molar

teeth are anchitherioid in pattern。



But the most important discovery of all is the Orohippus;

which comes from the Eocene formation; and is the oldest member

of the equine series; as yet known。 Here we find four complete

toes on the front limb; three toes on the hind limb; a well…

developed ulna; a well…developed fibula; and short…crowned

grinders of simple pattern。



Thus; thanks to these important researches; it has become

evident that; so far as our present knowledge extends; the

history of the horse…type is exactly and precisely that which

could have been predicted from a knowledge of the principles of

evolution。 And the knowledge we now possess justifies us

completely in the anticipation; that when the still lower Eocene

deposits; and those which belong to the Cretaceous epoch; have

yielded up their remains of ancestral equine animals; we shall

find; first; a form with four complete toes and a rudiment of

the innermost or first digit in front; with; probably; a

rudiment of the fifth digit in the hind foot; while; in still

older forms; the series of the digits will be more and more

complete; until we come to the five…toed animals; in which; if

the doctrine of evolution is well founded; the whole series must

have taken its origin。



That is what I mean by demonstrative evidence of evolution。

An inductive hypothesis is said to be demonstrated when the

facts are shown to be in entire accordance with it。 If that is

not scientific proof; there are no merely inductive conclusions

which can be said to be proved。 And the doctrine of evolution;

at the present time; rests upon exactly as secure a foundation

as the Copernican theory of the motions of the heavenly bodies

did at the time of its promulgation。 Its logical basis is

precisely of the same characterthe coincidence of the observed

facts with theoretical requirements。



The only way of escape; if it be a way of escape; from the

conclusions which I have just indicated; is the supposition that

all these different equine forms have been created separately at

separate epochs of time; and; I repeat; that of such an

hypothesis as this there neither is; nor can be; any scientific

evidence; and; assuredly; so far as I know; there is none which

is supported; or pretends to be supported; by evidence or

authority of any other kind。 I can but think that the time will

come when such suggestions as these; such obvious attempts to

escape the force of demonstration; will be put upon the same

footing as the supposition made by some writers; who are I

believe not completely extinct at present; that fossils are mere

simulacra; are no indications of the former existence of the

animals to which they seem to belong; but that they are either

sports of nature; or special creations; intendedas I heard

suggested the other dayto test our faith。



In fact; the whole evidence is in favour of evolution; and there

is none against it。 And I say this; although perfectly well

aware of the seeming difficulties which have been built up upon

what appears to the uninformed to be a solid foundation。 I meet

constantly with the argument that the doctrine of evolution

cannot be well founded; because it requires the lapse of a very

vast period of time; while the duration of life upon the earth

thus implied is inconsistent with the conclusions arrived at by

the astronomer and the physicist。 I may venture to say that I am

familiar with those conclusions; inasmuch as some years ago;

when President of the Geological Society of London; I took the

liberty of criticising them; and of showing in what respects; as

it appeared to me; they lacked complete and thorough

demonstration。 But; putting that point aside; suppose that; as

the astronomers; or some of them; and some physical

philosophers; tell us; it is impossible that life could have

endured upon the earth for as long a period as is required by

the doctrine of evolutionsupposing that to be provedI desire

to be informed; what is the foundation for the statement that

evolution does require so great a time? The biologist knows

nothing whatever of the amount of time which may be required for

the process of evolution。 It is a matter of fact that the equine

forms which I have described to you occur; in the order stated;

in the Tertiary formations。 But I have not the slightest means

of guessing whether it took a million of years; or ten millions;

or a hundred millions; or a thousand millions of years; to give

rise to that series of changes。 A biologist has no means of

arriving at any conclusion as to the amount of time which may be

needed for a certain quantity of organic change。 He takes his

time from the geologist。 The geologist; considering the rate at

which deposits are formed and the rate at which denudation goes

on upon the surface of the earth; arrives at more or less

justifiable conclusions as to the time which is required for the

deposit of a certain thickness of rocks; and if he tells me that

the Tertiary formations required 500;000;000 years for their

deposit; I suppose he has good ground for what he says; and I

take that as a measure of the duration of the evolution of the

horse from the Orohippus up to its present condition。

And; if he is right; undoubtedly evolution is a very slow

process; and requires a great deal of time。 But suppose; now;

that an astronomer or a physicistfor instance; my friend Sir

William Thomsontells me that my geological authority is quite

wrong; and that he has weighty evidence to show that life could

not possibly have existed upon the surface of the earth

500;000;000 years ago; because the earth would have then been

too hot to allow of life; my reply is: 〃That is not my affair;

settle that with the geologist; and when you have come to an

agreement among yourselves I will adopt your conclusion。〃

We take our time from the geologists and physicists; and it is

monstrous that; having taken our time from the physical

philosopher's clock; the physical philosopher should turn round

upon us; and say we are too fast or too slow。 What we desire to

know is; is it a fact that evolution took place? As to the

amount of time which evolution may have occupied; we are in the

hands of the physicist and the astronomer; whose business it is

to deal with those questions。



I have now; ladies and gentlemen; arrived at the conclusion of

the task which I set before myself when I undertook to deliver

these lectures。 My purpose has been; not to enable those among

you who have paid no attention to these subjects before; to

leave this room in a condition to decide upon the validity or

the invalidity of the hypothesis of evolution; but I have

desired to put before you the principles upon which all

hypotheses respecting the history of Nature must be judged;

and furthermore; to make apparent the nature of the evidence and

the amount of cogency which is to be expected and may be

obtained from it。 To this end; I have not hesitated to regard

you as genuine st

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