lectures on evolution-第15章
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peculiarities of that formpeculiarities which tend to show
that the European Hipparion is rather a member of a
collateral branch; than a form in the direct line of succession。
Next; in the backward order in time; is the Miohippus;
which corresponds pretty nearly with the Anchitherium of
Europe。 It presents three complete toesone large median and
two smaller lateral ones; and there is a rudiment of that digit;
which answers to the little finger of the human hand。
The European record of the pedigree of the horse stops here;
in the American Tertiaries; on the contrary; the series of
ancestral equine forms is continued into the Eocene formations。
An older Miocene form; termed Mesohippus; has three toes
in front; with a large splint…like rudiment representing the
little finger; and three toes behind。 The radius and ulna; the
tibia and the fibula; are distinct; and the short crowned molar
teeth are anchitherioid in pattern。
But the most important discovery of all is the Orohippus;
which comes from the Eocene formation; and is the oldest member
of the equine series; as yet known。 Here we find four complete
toes on the front limb; three toes on the hind limb; a well…
developed ulna; a well…developed fibula; and short…crowned
grinders of simple pattern。
Thus; thanks to these important researches; it has become
evident that; so far as our present knowledge extends; the
history of the horse…type is exactly and precisely that which
could have been predicted from a knowledge of the principles of
evolution。 And the knowledge we now possess justifies us
completely in the anticipation; that when the still lower Eocene
deposits; and those which belong to the Cretaceous epoch; have
yielded up their remains of ancestral equine animals; we shall
find; first; a form with four complete toes and a rudiment of
the innermost or first digit in front; with; probably; a
rudiment of the fifth digit in the hind foot; while; in still
older forms; the series of the digits will be more and more
complete; until we come to the five…toed animals; in which; if
the doctrine of evolution is well founded; the whole series must
have taken its origin。
That is what I mean by demonstrative evidence of evolution。
An inductive hypothesis is said to be demonstrated when the
facts are shown to be in entire accordance with it。 If that is
not scientific proof; there are no merely inductive conclusions
which can be said to be proved。 And the doctrine of evolution;
at the present time; rests upon exactly as secure a foundation
as the Copernican theory of the motions of the heavenly bodies
did at the time of its promulgation。 Its logical basis is
precisely of the same characterthe coincidence of the observed
facts with theoretical requirements。
The only way of escape; if it be a way of escape; from the
conclusions which I have just indicated; is the supposition that
all these different equine forms have been created separately at
separate epochs of time; and; I repeat; that of such an
hypothesis as this there neither is; nor can be; any scientific
evidence; and; assuredly; so far as I know; there is none which
is supported; or pretends to be supported; by evidence or
authority of any other kind。 I can but think that the time will
come when such suggestions as these; such obvious attempts to
escape the force of demonstration; will be put upon the same
footing as the supposition made by some writers; who are I
believe not completely extinct at present; that fossils are mere
simulacra; are no indications of the former existence of the
animals to which they seem to belong; but that they are either
sports of nature; or special creations; intendedas I heard
suggested the other dayto test our faith。
In fact; the whole evidence is in favour of evolution; and there
is none against it。 And I say this; although perfectly well
aware of the seeming difficulties which have been built up upon
what appears to the uninformed to be a solid foundation。 I meet
constantly with the argument that the doctrine of evolution
cannot be well founded; because it requires the lapse of a very
vast period of time; while the duration of life upon the earth
thus implied is inconsistent with the conclusions arrived at by
the astronomer and the physicist。 I may venture to say that I am
familiar with those conclusions; inasmuch as some years ago;
when President of the Geological Society of London; I took the
liberty of criticising them; and of showing in what respects; as
it appeared to me; they lacked complete and thorough
demonstration。 But; putting that point aside; suppose that; as
the astronomers; or some of them; and some physical
philosophers; tell us; it is impossible that life could have
endured upon the earth for as long a period as is required by
the doctrine of evolutionsupposing that to be provedI desire
to be informed; what is the foundation for the statement that
evolution does require so great a time? The biologist knows
nothing whatever of the amount of time which may be required for
the process of evolution。 It is a matter of fact that the equine
forms which I have described to you occur; in the order stated;
in the Tertiary formations。 But I have not the slightest means
of guessing whether it took a million of years; or ten millions;
or a hundred millions; or a thousand millions of years; to give
rise to that series of changes。 A biologist has no means of
arriving at any conclusion as to the amount of time which may be
needed for a certain quantity of organic change。 He takes his
time from the geologist。 The geologist; considering the rate at
which deposits are formed and the rate at which denudation goes
on upon the surface of the earth; arrives at more or less
justifiable conclusions as to the time which is required for the
deposit of a certain thickness of rocks; and if he tells me that
the Tertiary formations required 500;000;000 years for their
deposit; I suppose he has good ground for what he says; and I
take that as a measure of the duration of the evolution of the
horse from the Orohippus up to its present condition。
And; if he is right; undoubtedly evolution is a very slow
process; and requires a great deal of time。 But suppose; now;
that an astronomer or a physicistfor instance; my friend Sir
William Thomsontells me that my geological authority is quite
wrong; and that he has weighty evidence to show that life could
not possibly have existed upon the surface of the earth
500;000;000 years ago; because the earth would have then been
too hot to allow of life; my reply is: 〃That is not my affair;
settle that with the geologist; and when you have come to an
agreement among yourselves I will adopt your conclusion。〃
We take our time from the geologists and physicists; and it is
monstrous that; having taken our time from the physical
philosopher's clock; the physical philosopher should turn round
upon us; and say we are too fast or too slow。 What we desire to
know is; is it a fact that evolution took place? As to the
amount of time which evolution may have occupied; we are in the
hands of the physicist and the astronomer; whose business it is
to deal with those questions。
I have now; ladies and gentlemen; arrived at the conclusion of
the task which I set before myself when I undertook to deliver
these lectures。 My purpose has been; not to enable those among
you who have paid no attention to these subjects before; to
leave this room in a condition to decide upon the validity or
the invalidity of the hypothesis of evolution; but I have
desired to put before you the principles upon which all
hypotheses respecting the history of Nature must be judged;
and furthermore; to make apparent the nature of the evidence and
the amount of cogency which is to be expected and may be
obtained from it。 To this end; I have not hesitated to regard
you as genuine st